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Haplodrassus signifer (C. L. Koch, 1839)

  FAMILY: GNAPHOSIDAE Pocock, 1898
    Genus: Haplodrassus Chamberlin, 1922
        Species: Haplodrassus signifer (C. L. Koch, 1839) LSID
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Author: Shorthouse, David P. Biography

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Common Names

Family Common Name:  stealthy ground spiders
Genus Common Name:  none
Species Common Name:  none

Distribution

Global Distribution: Holarctic
Nearctic States & Provinces:   AB ON CO MT NH OK PA SC WY
Collection Locales Mapped by Year Collected: 

Haplodrassus signifer (C. L. Koch, 1839) collection map
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Specimen Search

Specimen list
73, 65

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GeoRSS

Natural History, Phenology & Image(s)

Feeding Guild: ground runners
Feeding guild when noted defined by Uetz, G. W., J. Halaj, and A. B. Cady. 1999. Guild structure of spiders in major crops. Journal of Arachnology 27:270-280. PDF
Recorded Habitats: under stones, logs, ground debris, grassland and wheat fields, salt marshes, deciduous and coniferous forests, in association with ephedra, sagebrush, and sphagnum (Platnick & Dondale, 1992).
Life & Natural History: (not yet recorded or unknown)

Seasonality of Specimen Records:


Image(s) of Habitus: Submit an image email an image

Details About Males

Description: (not yet recorded or unknown)

Male Dimensions:
Male dimensionsA = 2.9 mm 
A+B = 6.51 mm 
C = 2.26 mm 
Image of External Male Genitalia:

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Details About Females

Description: (not yet recorded or unknown)

Female Dimensions:
Female dimensions A = 2.84 mm 
A+B = 7.18 mm 
C = 2.2 mm 
Image of Epigynum:

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Descriptions Source:
Platnick, N. I. & C. D. Dondale. 1992. The insects and arachnids of Canada, Part 19. The ground spiders of Canada and Alaska (Araneae: Gnaphosidae). Research Branch, Agriculture Canada, Publ. 1875: 1-297.

Synonyms and Chresonyms

Drassus signifer C. L. Koch 1839 urn:lsid:ubio.org:namebank:2094443
Drassus troglodytes C. L. Koch 1839 urn:lsid:ubio.org:namebank:2094444
Haplodrassus signifer C. L. Koch 1839 urn:lsid:ubio.org:namebank:9737968
Clubiona troglodytes Walckenaer 1842 urn:lsid:ubio.org:namebank:2094445
Drassus clavator O. P.-Cambridge 1860 urn:lsid:ubio.org:namebank:2094446
Drassus clavator Blackwall 1861 urn:lsid:ubio.org:namebank:2094447
Drassus troglodytes L. Koch 1866 urn:lsid:ubio.org:namebank:2094448
Drassus troglodytes Menge 1875 urn:lsid:ubio.org:namebank:2094450
Drassus robustus Emerton 1890 urn:lsid:ubio.org:namebank:2094451
Drassus mysticus O. P.-Cambridge 1894 urn:lsid:ubio.org:namebank:2094452
Teminius nigriceps Banks 1895 urn:lsid:ubio.org:namebank:2094453
Drassus placidus Banks 1896 urn:lsid:ubio.org:namebank:2094454
Drassus troglodytes Becker 1896 urn:lsid:ubio.org:namebank:2094455
Drassus troglodytes Chyzer & Kulczynski 1897 urn:lsid:ubio.org:namebank:2094456
Drassodes ferrum-equinum F. O. P.-Cambridge 1899 urn:lsid:ubio.org:namebank:2094457
Prosthesima decepta Banks 1900 urn:lsid:ubio.org:namebank:2094458
Drassus troglodytes Bösenberg 1902 urn:lsid:ubio.org:namebank:2094459
Drassodes robustus Bryant 1908 urn:lsid:ubio.org:namebank:2094461
Drassodes troglodytes Lessert 1910 urn:lsid:ubio.org:namebank:2094462
Syrisca nigriceps Petrunkevitch 1911 urn:lsid:ubio.org:namebank:2094464
Zelotes decepta Petrunkevitch 1911 urn:lsid:ubio.org:namebank:2094463
Drassodes signifer Comstock 1912 urn:lsid:ubio.org:namebank:2094465
Drassodes troglodytes Simon 1914 urn:lsid:ubio.org:namebank:2094466
Drassodes beaufortensis Strand 1916 urn:lsid:ubio.org:namebank:2094467
Haplodrassus signifer Chamberlin 1922 urn:lsid:ubio.org:namebank:2094468
Haplodrassus signifer Reimoser 1937 urn:lsid:ubio.org:namebank:2094469
Haplodrassus dystactus Chamberlin & Gertsch 1940 urn:lsid:ubio.org:namebank:2094470
Haplodrassus signifer Comstock 1940 urn:lsid:ubio.org:namebank:2094472
Haplodrassus signifer Palmgren 1943 urn:lsid:ubio.org:namebank:2094473
Drassodes signifer Braendegaard 1946 urn:lsid:ubio.org:namebank:2094475
Haplodrassus signifer Tullgren 1946 urn:lsid:ubio.org:namebank:2094474
Haplodrassus signifer Kaston 1948 urn:lsid:ubio.org:namebank:2094476
Drassodes signifer Locket & Millidge 1951 urn:lsid:ubio.org:namebank:2094477
Haplodrassus signifer Braendegaard 1966 urn:lsid:ubio.org:namebank:2094478
Haplodrassus signifer Miller 1971 urn:lsid:ubio.org:namebank:2094480
Haplodrassus signifer Tyschchenko 1971 urn:lsid:ubio.org:namebank:2094479
Haplodrassus signifer Platnick & Shadab 1975 urn:lsid:ubio.org:namebank:2094481
Haplodrassus signifer Miller & Buchar 1977 urn:lsid:ubio.org:namebank:2094483
Haplodrassus signifer Paik 1978 urn:lsid:ubio.org:namebank:2094484
Drassodes signifer Jia & Zhu 1983 urn:lsid:ubio.org:namebank:2094485
Haplodrassus signifer Thaler 1984 urn:lsid:ubio.org:namebank:2094486
Haplodrassus signifer Grimm 1985 urn:lsid:ubio.org:namebank:2094488
Haplodrassus signifer Roberts 1985 urn:lsid:ubio.org:namebank:2094489
Haplodrassus signifer Heiss & Allen 1986 urn:lsid:ubio.org:namebank:2094490
Haplodrassus aeneus Hu & Li 1987 urn:lsid:ubio.org:namebank:2094247
Haplodrassus signifer Izmailova 1989 urn:lsid:ubio.org:namebank:2094491
Haplodrassus signifer Heimer & Nentwig 1991 urn:lsid:ubio.org:namebank:2094492
Haplodrassus signifer Noordam 1992 urn:lsid:ubio.org:namebank:2094496
Haplodrassus signifer Paik 1992 urn:lsid:ubio.org:namebank:2094495
Haplodrassus signifer Platnick & Dondale 1992 urn:lsid:ubio.org:namebank:2094494
Haplodrassus signifer Roberts 1995 urn:lsid:ubio.org:namebank:2094497
Haplodrassus signifer Agnarsson 1996 urn:lsid:ubio.org:namebank:2094500
Haplodrassus signifer Marusik, Hippa & Koponen 1996 urn:lsid:ubio.org:namebank:2094498
Haplodrassus signifer Ovtsharenko & Marusik 1996 urn:lsid:ubio.org:namebank:2094499
Haplodrassus signifer Bellmann 1997 urn:lsid:ubio.org:namebank:2094502
Haplodrassus signifer Mcheidze 1997 urn:lsid:ubio.org:namebank:2094501
Haplodrassus signifer Roberts 1998 urn:lsid:ubio.org:namebank:2094503
Drassodes signifer Hu 2001 urn:lsid:ubio.org:namebank:2031430
Haplodrassus aeneus Hu 2001 urn:lsid:ubio.org:namebank:2031340
Haplodrassus signifer Chatzaki, Thaler & Mylonas 2002 urn:lsid:ubio.org:namebank:9734110
Haplodrassus signifer Paquin & Dupérré 2003 urn:lsid:ubio.org:namebank:2010532
Haplodrassus signifer Levy 2004 urn:lsid:ubio.org:namebank:9738283
Haplodrassus signifer Song, Zhu & Zhang 2004 urn:lsid:ubio.org:namebank:9738284
Haplodrassus signifer Almquist 2006

Taxonomic References Instructions

Agnarsson, I. 1996. Íslenskar köngulaer. Fjölrit Nàttùrufraedistofnunar 31: 1-175.

Almquist, S. 2006. Swedish Araneae, part 2--families Dictynidae to Salticidae. Insect Syst. Ecol., Suppl. 63: 285-601.

Banks, N. 1900. Some Arachnida from Alabama. Proc. Acad. nat. Sci. Philad. 52: 529-543.

Banks, N. 1896. New North American spiders and mites. Trans. Amer. ent. Soc. 23: 57-77.

Banks, N. 1895. The Arachnida of Colorado. Ann. New York Acad. Sci. 8: 415-434.

Becker, L. 1896. Les arachnides de Belgique. Ann. Mus. Roy. Hist. nat. Belg. 12: 1-3788.

Bellmann, H. 1997. Kosmos-Atlas Spinnentiere Europas. Frankh-Kosmos Verlag, Stuttgart, 304 pp.

Blackwall, J. 1861. A history of the spiders of Great Britain and Ireland. London, 1: 1-174.

Bösenberg, W. 1902. Die Spinnen Deutschlands. II-IV. Zoologica (Stuttgart) 14: 97-384.

Braendegaard, J. 1966. Edderkopper: Eller Spindlere I. Danmarks Fauna 72: 1-224.

Braendegaard, J. 1946. The spiders (Araneina) of East Greenland: A faunistic and zoogeographical investigation. Meddr Grønland 121(15): 1-128.

Bryant, E. B. 1908. List of the Araneina. In Fauna of New England, 9. Occas. Pap. Boston Soc. nat. Hist. 7: 1-105.

Cambridge, F. O. P.-. 1899. Arachnida - Araneida and Opiliones. In Biologia Centrali-Americana, Zoology. London, 2: 41-88.

Cambridge, O. P.-. 1894. On new and rare British spiders found in 1893; with rectifications of synonyms. Proc. Dorset nat. Hist Field Club 15: 103-116.

Cambridge, O. P.-. 1860. Descriptions of two British spiders new to science. Ann. Mag. nat. Hist. (3) 5: 171-174.

Chamberlin, R. V. 1922. The North American spiders of the family Gnaphosidae. Proc. biol. Soc. Wash. 35: 145-172.

Chamberlin, R. V. & W. J. Gertsch. 1940. Descriptions of new Gnaphosidae from the United States. Amer. Mus. Novit. 1068: 1-19.

Chatzaki, M., K. Thaler & M. Mylonas. 2002. Ground spiders (Gnaphosidae; Araneae) of Crete (Greece). Taxonomy and distribution. I. Rev. suisse Zool. 109: 559-601.

Chyzer, C. & W. Kulczynski. 1897. Araneae hungariae. Budapest, 2: 151-366.

Comstock, J. H. 1940. The spider book, revised and edited by W. J. Gertsch. Cornell Univ. Press, Ithaca, xi + 727 pp.

Comstock, J. H. 1912. The spider book; a manual for the study of the spiders and their near relatives, the scorpions, pseudoscorpions, whipscorpions, harvestmen and other members of the class Arachnida, found in America north of Mexico, with analytical keys for their classification and popular accounts of their habits. Garden City, New York, pp. 1-721

Emerton, J. H. 1890. New England spiders of the families Drassidae, Agalenidae and Dysderidae. Trans. Connect. Acad. Arts Sci. 8: 166-206.

Grimm, U. 1985. Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abh. naturw. Ver. Hamb. 26: 1-318.

Heimer, S. & W. Nentwig. 1991. Spinnen Mitteleuropas: Ein Bestimmungsbuch. Verlag Paul Parey, Berlin, 543 pp.

Heiss, J. S. & R. T. Allen. 1986. The Gnaphosidae of Arkansas. Bull. Ark. agric. Exp. Stn 885: 1-67.

Hu, J. L. 2001. Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, 658 pp.

Izmailova, M. V. 1989. [Fauna of Spiders of South Part of Eastern Siberia]. Irkutsk, State Univ. Publ., 184 pp.

Jia, L. D. & C. D. Zhu. 1983. [New record of spider of the genus Drassodes from Hui Autonomous Region of Ningxia, China (Araneae: Gnaphosidae)]. J. Bethune med. Univ. 9(suppl.): 167.

Kaston, B. J. 1948. Spiders of Connecticut. Bull. Conn. St. geol. nat. Hist. Surv. 70: 1-874.

Koch, C. L. 1839. Die Arachniden. Nürnberg, Funfter Band, pp. 125-158, Sechster Band, pp. 1-156, Siebenter Band, pp. 1-106.

Koch, L. 1866. Die Arachniden-Familie der Drassiden. Nürnberg, Hefte 1-6, pp. 1-304.

Lessert, R. de. 1910. Catalogue des invertebres de la Suisse. Fasc. 3, Araignées. Musée d'histoire naturelle de Genève, pp. 1-635.

Levy, G. 2004. Spiders of the genera Drassodes and Haplodrassus (Araneae, Gnaphosidae) from Israel. Israel J. Zool. 50: 1-37. To publisher...

Locket, G. H. & A. F. Millidge. 1951. British spiders. Ray Society, London, 1: 1-310.

Marusik, Y. M., H. Hippa & S. Koponen. 1996. Spiders (Araneae) from the Altai area, southern Siberia. Acta Zool. Fennica 201: 11-45.

Mcheidze, T. S. 1997. [Spiders of Georgia: Systematics, Ecology, Zoogeographic Review]. Tbilisi Univ., 390 pp. (in Georgian).

Menge, A. 1875. Preussische Spinnen. VII. Abtheilung. Schrift. naturf. Ges. Danzig (N. F.) 3: 375-422.

Miller, F. 1971. Pavouci-Araneida. Klíc zvíreny CSSR 4: 51-306.

Miller, F. & J. Buchar. 1977. Neue Spinnenarten aus der Gattung Zelotes Distel [sic] und Haplodrassus Chamberlin (Araneae, Gnaphosidae). Acta Univ. Carol. (Biol.) 1974: 157-171.

Noordam, A. P. 1992. Gnaphosidae (Araneae) of the Netherlands, drawings, some notes, and some thoughts about ant-mimicry. Published by the author, Groenesteeg 104, 2312 SR Leiden, Netherlands.

Ovtsharenko, V. I. & Y. M. Marusik. 1996. Additional data on the spiders of the family Gnaphosidae (Aranei) of the north-east of Asia. In ntomological Studies in the North-East of the USSR. USSR Academy of Sciences, Institute of Biological Problems of the North, Vladivostok, pp. 114-130 (

Paik, K. Y. 1992. Korean spiders of the genus Haplodrassus (Araneae: Gnaphosidae) II. Korean Arachnol. 8: 85-93.

Paik, K. Y. 1978. Araneae. Illustr. Fauna Flora Korea 21: 1-548.

Palmgren, P. 1943. Die Spinnenfauna Finnlands II. Act. zool. Fennica 36: 1-115.

Paquin, P. & N. Dupérré. 2003. Guide d'identification des araignées de Québec. Fabreries, Suppl. 11 1-251.

Petrunkevitch, A. 1911. A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greeland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bull. Amer. Mus. Nat. Hist. 29: 1-791. To publisher...

Platnick, N. I. & C. D. Dondale. 1992. The insects and arachnids of Canada, Part 19. The ground spiders of Canada and Alaska (Araneae: Gnaphosidae). Research Branch, Agriculture Canada, Publ. 1875: 1-297.

Platnick, N. I. & M. U. Shadab. 1975. A revision of the spider genera Haplodrassus and Orodrassus (Araneae, Gnaphosidae) in North America. Am. Mus. Novit. 2583: 1-40. To publisher...

Reimoser, E. 1937. Spinnentiere oder Arachnoidea. VII. Gnaphosidae oder Plattbauchspinnen. In Die Tierwelt Deutschlands. Jena, 33(16): 1-41.

Roberts, M. J. 1998. Spinnengids. Tirion, Baarn, Netherlands, 397 pp.

Roberts, M. J. 1995. Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp.

Roberts, M. J. 1985. The spiders of Great Britain and Ireland, Volume 1: Atypidae to Theridiosomatidae. Harley Books, Colchester, England.

Simon, E. 1914. Les arachnides de France. Synopsis générale et catalogue des espèces françaises de l'ordre des Araneae; 1re partie. Paris, 6: 1-308.

Song, D. X., M. S. Zhu & F. Zhang. 2004. Fauna Sinica: Invertebrata Vol. 39: Arachnida: Araneae: Gnaphosidae. Science Press, Beijing, ix + 362 pp.

Strand, E. 1916. Systematische-faunistiche Studien über paläarktische, afrikanische und amerikanische Spinnen des Senckenbergischen Museums. Arch. Naturg. 81(A9): 1-153.

Thaler, K. 1984. Haplodrassus aenus n. sp. aus Österreich und der Schweiz (Arachnida: Araneae, Gnaphosidae). Mitt. schweiz. ent. Ges. 57: 189-193.

Tullgren, A. 1946. Svenska spindelfauna: 3. Egentliga spindlar. Araneae. Fam. 5-7. Clubionidae, Zoridae och Gnaphosidae. Entomologiska Föreningen, Stockholm, 141 pp.

Tyschchenko, V. P. 1971. Opredelitel' paukov evropejskoj casti SSSR. Leningrad, pp. 1-281.

Walckenaer, C. A. 1842. Histoire naturelle des Insects. Aptères. Paris, 2: 1-549.

Nomenclature and taxonomic references from the World Spider Catalog

Natural History References Instructions

Platnick, N. I. & C. D. Dondale. 1992. The insects and arachnids of Canada, Part 19. The ground spiders of Canada and Alaska (Araneae: Gnaphosidae). Research Branch, Agriculture Canada, Publ. 1875: 1-297.

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Author of this page in progress: Shorthouse, David P. dshorthouse(at)eol.org
Text Last Modified: 2006-03-15T18:32:13Z